Andy Reynolds

NameAndy Reynolds
Job titleEmeritus
Email addressandy.reynolds@rothamsted.ac.uk
DepartmentIntelligent Data Ecosystems
Research clusterCAS: Biological Physics and Engineering
Preferred citationReynolds, A. M.
OfficeHarpenden

Research outputs

Mosquito swarms shear harden

Reynolds, A. M. 2023. Mosquito swarms shear harden. European Physical Journal E. 46, p. 126. https://doi.org/10.1140/epje/s10189-023-00379-3

Phase transitions in insect swarms

Reynolds, A. M. 2023. Phase transitions in insect swarms . Physical Biology. 20 (5), p. 054001. https://doi.org/10.1088/1478-3975/aceece

A sublethal concentration of Sulfoxaflor has minimal impact on buff-tailed bumblebee (Bombus terrestris) locomotor behaviour under aversive conditioning

James, L., Reynolds, A. M., Mellor, I. R. and Davies, T. G. E. 2023. A sublethal concentration of Sulfoxaflor has minimal impact on buff-tailed bumblebee (Bombus terrestris) locomotor behaviour under aversive conditioning. Toxics. 11 (3), p. 279. https://doi.org/10.3390/toxics11030279

Stochasticity may generate coherent motion in bird flocks

Reynolds, A. M. 2023. Stochasticity may generate coherent motion in bird flocks . Physical Biology. 20 (2), p. 025002. https://doi.org/10.1088/1478-3975/acbad7

Swarm formation as backward diffusion

Reynolds, A. M. and Ouellette, N. T. 2023. Swarm formation as backward diffusion. Physical Biology. 20 (2), p. 026002. https://doi.org/10.1088/1478-3975/acb986

Spaces between insects in laboratory swarms move like insects in natural swarms

Reynolds, A. M. 2023. Spaces between insects in laboratory swarms move like insects in natural swarms. EPL. 141 (1), p. 17001. https://doi.org/10.1209/0295-5075/acab7c

Comment on ‘A physics perspective on collective animal behavior’ 2022 Phys. Biol. 19 021004

Reynolds, A. M. 2022. Comment on ‘A physics perspective on collective animal behavior’ 2022 Phys. Biol. 19 021004. Physical Biology. 19 (6), p. 068001. https://doi.org/10.1088/1478-3975/ac8fd5

Stochastic modelling of bird flocks – accounting for the cohesiveness of collective motion

Reynolds, A. M., McIvor, G. E., Thornton, A., Yang, P. and Ouellette, N. T. 2022. Stochastic modelling of bird flocks – accounting for the cohesiveness of collective motion. Journal of the Royal Society Interface. 19, p. 20210745. https://doi.org/10.1098/rsif.2021.0745

Understanding the thermodynamic properties of insect swarms

Reynolds, A. M. 2021. Understanding the thermodynamic properties of insect swarms. Scientific Reports. 11, p. 14979. https://doi.org/10.1038/s41598-021-94582-x

Weierstrassian Levy walks are a by-product of crawling

Reynolds, A. M. 2021. Weierstrassian Levy walks are a by-product of crawling. European Physical Journal E. 44, p. 96. https://doi.org/10.1140/epje/s10189-021-00100-2

Loss of mechanical stiffness and the emergence of Lévy walks in active T cells

Reynolds, A. M. 2021. Loss of mechanical stiffness and the emergence of Lévy walks in active T cells. Physics Letters A. 408 (27 Aug), p. 127507. https://doi.org/10.1016/j.physleta.2021.127507

Harmonic radar tracking reveals that honeybee drones navigate between multiple aerial leks

Woodgate, J. L., Makinson, J. C., Rossi, N., Reynolds, A. M., Rawlings, C. J. and Chittka, L. 2021. Harmonic radar tracking reveals that honeybee drones navigate between multiple aerial leks. iScience. 24 (6), p. 102499. https://doi.org/10.1016/j.isci.2021.102499

Scale-free movement patterns in termites emerge from social interactions and preferential attachments

Paiva, L. R., Marins, A., Cristaldo, P. F., Ribeiro, D., Alves, S. G., Reynolds, A. M., DeSouza, O. and Miramontes, O. 2021. Scale-free movement patterns in termites emerge from social interactions and preferential attachments. Proceedings of the National Academy of Sciences. 118 (20), p. e2004369118. https://doi.org/10.1073/pnas.2004369118

An equation of state for insect swarms

Sinhuber, M., Van der Vaart, K., Yenchia, F., Reynolds, A. M. and Ouellette, N.T. 2021. An equation of state for insect swarms. Scientific Reports. 11 (3773). https://doi.org/10.1038/s41598-021-83303-z

Intrinsic stochasticity and the emergence of collective behaviours in insect swarms

Reynolds, A. M. 2020. Intrinsic stochasticity and the emergence of collective behaviours in insect swarms. European Physical Journal E. 44 (2), p. 22. https://doi.org/10.1140/epje/s10189-021-00040-x

Insect swarms can be bound together by repulsive forces

Reynolds, A. M. 2020. Insect swarms can be bound together by repulsive forces. European Physical Journal E. 43, p. 39. https://doi.org/10.1140/epje/i2020-11963-x

Environmental Perturbations Induce Correlations in Midge Swarms

Van Der Vaart, K., Sinhuber, M., Reynolds, A. M. and Ouellette, N. T. 2020. Environmental Perturbations Induce Correlations in Midge Swarms. Journal of the Royal Society Interface. 17 (164), p. 20200018. https://doi.org/10.1098/rsif.2020.0018

Similarities between Insect Swarms and Isothermal Globular Clusters

Gorbonos, D., Van der Vaart, K., Sinhuber, M., Puckett, J. G., Ouellette, N. T., Reynolds, A. M. and Gov, N. S. 2020. Similarities between Insect Swarms and Isothermal Globular Clusters. Physical Review Research. 2 (1), p. 013271. https://doi.org/10.1103/PhysRevResearch.2.013271

Linking small-scale flight manoeuvers and density profiles to the vertical movement of insects in the nocturnal stable boundary layer

Wainwright, C. E., Reynolds, D. R. and Reynolds, A. M. 2020. Linking small-scale flight manoeuvers and density profiles to the vertical movement of insects in the nocturnal stable boundary layer. Scientific Reports. 10, p. 1019. https://doi.org/10.1038/s41598-020-57779-0

Do bumblebees have signatures? Demonstrating the existence of a speed-curvature power law in Bombus terrestris locomotion patterns.

James, L., Davies, T. G. E., Lim, K. S. and Reynolds, A. M. 2020. Do bumblebees have signatures? Demonstrating the existence of a speed-curvature power law in Bombus terrestris locomotion patterns. PLOS ONE. 15 (1), p. e0226393. https://doi.org/10.1371/journal.pone.0226393

On the emergence of gravitational-like forces in insect swarms

Reynolds, A. M. 2019. On the emergence of gravitational-like forces in insect swarms. Journal of the Royal Society Interface. 16 (160), p. 20190404. https://doi.org/10.1098/rsif.2019.0404

Investigating vertical motion of small insects in the atmospheric boundary layer using millimetre-wavelength radar and Doppler LIDAR

Wainwright, C. E., Stepanian, P. M., Reynolds, D. R. and Reynolds, A. M. 2019. Investigating vertical motion of small insects in the atmospheric boundary layer using millimetre-wavelength radar and Doppler LIDAR. Institution of Engineering and Technology. https://doi.org/10.1049/joe.2019.0596

Mechanical spectroscopy of insect swarms

Van der Vaart, K., Sinhuber, M., Reynolds, A. M. and Ouellette, N.T. 2019. Mechanical spectroscopy of insect swarms. Science Advances. 5 (7), p. eaaw9305. https://doi.org/10.1126/sciadv.aaw9305

On the origin of the tensile strength of insect swarms

Reynolds, A. M. 2019. On the origin of the tensile strength of insect swarms. Physical Biology. 16 (4), p. 046002. https://doi.org/10.1088/1478-3975/ab12b9

Harmonic radar tracking reveals random dispersal pattern of bumblebee (Bombus terrestris) queens after hibernation

Makinson, J. C., Woodgate, J. L., Reynolds, A. M., Capaldi, E. A., Perry, C. J. and Chittka, L. 2019. Harmonic radar tracking reveals random dispersal pattern of bumblebee (Bombus terrestris) queens after hibernation. Scientific Reports. 9 (4651). https://doi.org/10.1038/s41598-019-40355-6

Fluctuating environments drive insect swarms into a new state that is robust to perturbations

Reynolds, A. M. 2018. Fluctuating environments drive insect swarms into a new state that is robust to perturbations. EPL. 124, p. 38001. https://doi.org/10.1209/0295-5075/124/38001

Incorporating terminal velocities into Lagrangian stochastic models of particle dispersal in the atmospheric boundary layer

Reynolds, A. M. 2018. Incorporating terminal velocities into Lagrangian stochastic models of particle dispersal in the atmospheric boundary layer. Scientific Reports. 8, p. 16843. https://doi.org/10.1038/s41598-018-34924-4

Olfactory-cued navigation in shearwaters: linking movement patterns to mechanisms

Abolaffio, M., Reynolds, A. M., Cecere, J. G., Paiva, V. H. and Focardi, S. 2018. Olfactory-cued navigation in shearwaters: linking movement patterns to mechanisms. Scientific Reports. 8 (11590). https://doi.org/10.1038/s41598-018-29919-0

Passive particles Lévy walk through turbulence mirroring the diving patterns of marine predators

Reynolds, A. M. 2018. Passive particles Lévy walk through turbulence mirroring the diving patterns of marine predators. Journal of Physics Communications. 2 (8), p. 085003. https://doi.org/10.1088/2399-6528/aad498

Distinguishing between apparent and actual randomness: a preliminary examination with Australian ants

Reynolds, A. M., Ferdous, M. J. and Cheng, K. 2018. Distinguishing between apparent and actual randomness: a preliminary examination with Australian ants. Behavioral Ecology and Sociobiology. 72 (113). https://doi.org/10.1007/s00265-018-2527-1

Levy foraging patterns of rural humans

Reynolds, A. M., Ceccon, E., Baldauf, C., Medeiros, T. K. and Miramontes, O. 2018. Levy foraging patterns of rural humans. PLOS ONE. 13 (6), p. e0199099. https://doi.org/10.1371/journal.pone.0199099

More on midges

Reynolds, A. M. 2018. More on midges. Physics World. 31 (3), pp. 25-25. https://doi.org/10.1088/2058-7058/31

Current status and future directions of Levy walk research

Reynolds, A. M. 2018. Current status and future directions of Levy walk research. Biology Open. 7 (1), p. bio030106. https://doi.org/10.1242/bio.030106

Langevin dynamics encapsulate the microscopic and emergent macroscopic properties of midge swarms

Reynolds, A. M. 2018. Langevin dynamics encapsulate the microscopic and emergent macroscopic properties of midge swarms. Journal of the Royal Society Interface. 15. https://doi.org/10.1098/rsif.2017.0806

Continuous Radar Tracking Illustrates the Development of Multi-destination Routes of Bumblebees

Woodgate, J. L., Makinson, J. C., Lim, K. S., Reynolds, A. M. and Chittka, L. 2017. Continuous Radar Tracking Illustrates the Development of Multi-destination Routes of Bumblebees. Scientific Reports. 7 (1), p. 17323. https://doi.org/10.1038/s41598-017-17553-1

The movement of small insects in the convective boundary layer: linking patterns to processes

Wainwright, C. E., Stepanian, P. M., Reynolds, D. R. and Reynolds, A. M. 2017. The movement of small insects in the convective boundary layer: linking patterns to processes. Scientific Reports. 7 (1), p. 5438. https://doi.org/10.1038/s41598-017-04503-0

The Weierstrassian movement patterns of snails

Reynolds, A. M., Santini, G., Chelazzi, G. and Focardi, S. 2017. The Weierstrassian movement patterns of snails. Royal Society Open Science. 4 (6). https://doi.org/10.1098/rsos.160941

Chaotic model for lévy walks in swarming bacteria

Ariel, G., Be’er, A. and Reynolds, A. M. 2017. Chaotic model for lévy walks in swarming bacteria. Physical Review Letters. 118 (22), p. 228102. https://doi.org/10.1103/PhysRevLett.118.228102

Are midge swarms bound together by an effective velocity-dependent gravity?

Reynolds, A. M., Sinhuber, M. and Ouellette, N. T. 2017. Are midge swarms bound together by an effective velocity-dependent gravity? European Physical Journal E. 40 (4), p. 46. https://doi.org/10.1140/epje/i2017-11531-7

Optimal search patterns in honeybee orientation flights are robust against emerging infectious diseases

Wolf, S., Nicholls, E., Reynolds, A. M., Wells, P. M., Lim, K. S., Paxton, R. J. and Osborne, J. L. 2016. Optimal search patterns in honeybee orientation flights are robust against emerging infectious diseases. Scientific Reports. 6, p. 32612. https://doi.org/10.1038/srep32612

Orientation in high-flying migrant insects in relation to flows: mechanisms and strategies

Reynolds, A. M., Reynolds, D. R., Sane, S. P., Hu, G. and Chapman, J. W. 2016. Orientation in high-flying migrant insects in relation to flows: mechanisms and strategies. Philosophical Transactions of the Royal Society B-Biological Sciences. 371 (1704), p. 20150392. https://doi.org/10.1098/rstb.2015.0392

Life-long radar tracking of bumblebees

Woodgate, J. L., Makinson, J. C., Lim, K. S., Reynolds, A. M. and Chittka, L. 2016. Life-long radar tracking of bumblebees. PLOS ONE. 11 (8). https://doi.org/10.1371/journal.pone.0160333

Swarm dynamics may give rise to Lévy flights

Reynolds, A. M. and Ouellette, N. T. 2016. Swarm dynamics may give rise to Lévy flights. Scientific Reports. 6, p. 30515. https://doi.org/10.1038/srep30515

Signatures of active and passive optimized Levy searching in jellyfish

Reynolds, A. M. 2014. Signatures of active and passive optimized Levy searching in jellyfish. Journal of the Royal Society Interface. 11 (99), p. 20140665. https://doi.org/10.1098/rsif.2014.0665

The Cahn-Hilliard phase separation principle maybe the tip of an iceberg Comment on "Phase separation driven by density-dependent movement: A novel mechanism for ecological patterns" by Q.-X. Liu et al

Reynolds, A. M. 2016. The Cahn-Hilliard phase separation principle maybe the tip of an iceberg Comment on "Phase separation driven by density-dependent movement: A novel mechanism for ecological patterns" by Q.-X. Liu et al. Physics of Life Reviews. 19, pp. 135-136. https://doi.org/10.1016/j.plrev.2016.09.007

Wind-related orientation patterns in diurnal, crepuscular and nocturnal high-altitude insect migrants

Hu, G., Lim, K. S., Reynolds, D. R., Reynolds, A. M. and Chapman, J. W. 2016. Wind-related orientation patterns in diurnal, crepuscular and nocturnal high-altitude insect migrants. Frontiers in Behavioral Neuroscience. 10. https://doi.org/10.3389/fnbeh.2016.00032

Similarities and differences in path integration and search in two species of desert ants inhabiting a visually rich and a visually barren habitat

Schultheiss, P., Stannard, T., Pereira, S., Reynolds, A. M., Wehner, R. and Cheng, K. 2016. Similarities and differences in path integration and search in two species of desert ants inhabiting a visually rich and a visually barren habitat. Behavioral Ecology and Sociobiology. 70 (8), pp. 1319-1329. https://doi.org/10.1007/s00265-016-2140-0

How far can oak pollen be transported in the atmosphere in a single day?

Fernandez-Rodriguez, S., Maya-Manzano, J. M., Smith, M., Tormo-Molina, R., Reynolds, A. M., Silva-Palacios, I., Gonzalo-Garijo, A. and Sadys, M. 2016. How far can oak pollen be transported in the atmosphere in a single day? Proceedings 6th European Symposium in Aerobiology, 18-22 July, Lyon, France, 2016. pp. Poster 2407

Pollen dispersal in a hybrid cereal canopy

Sadys, M., West, J. S., Spiller, M., Melichar, G., Stiewe, G. and Reynolds, A. M. 2016. Pollen dispersal in a hybrid cereal canopy. In : Proceedings 6th European Symposium on Aerobiology, At Lyon, France, 18-22 July, 2016 . pp. Oral 2315

Airborne Quercus pollen in SW Spain: Identifying favourable conditions for atmospheric transport and potential source areas

Maya-Manzanoa, J. M., Fernandez-Rodriguez, S., Smith, M., Tormo-Molina, R., Reynolds, A. M., Silva-Palacios, I., Gonzalo-Garijo, A. and Sadys, M. 2016. Airborne Quercus pollen in SW Spain: Identifying favourable conditions for atmospheric transport and potential source areas. Science of the Total Environment. 571, pp. 1037-1047. https://doi.org/10.1016/j.scitotenv.2016.07.094

Signatures of a globally optimal searching strategy in the three-dimensional foraging flights of bumblebees

Lihoreau, M., Ings, T. C., Chittka, L. and Reynolds, A. M. 2016. Signatures of a globally optimal searching strategy in the three-dimensional foraging flights of bumblebees. Scientific Reports. 6, p. Article no 30401. https://doi.org/10.1038/srep30401

Signatures of chaos in animal search patterns

Reynolds, A. M., Bartumeus, F., Kolzch, A. and van de Koppel, J. 2016. Signatures of chaos in animal search patterns. Scientific Reports. 6, p. 23492. https://doi.org/10.1038/srep23492

Levy patterns in seabirds are multifaceted describing both spatial and temporal patterning

Reynolds, A. M., Paiva, V. H., Cecere. J. G. and Focardi, S. 2016. Levy patterns in seabirds are multifaceted describing both spatial and temporal patterning. Frontiers in Zoology. 13, p. 29. https://doi.org/10.1186/s12983-016-0160-2

Detection of flow direction in high-flying insect and songbird migrants

Chapman, J. W., Nilsson, C., Lim, K. S., Backman, J., Reynolds, D. R., Alerstam, T. and Reynolds, A. M. 2015. Detection of flow direction in high-flying insect and songbird migrants. Current Biology. 25, pp. R751-R752. https://doi.org/10.1016/j.cub.2015.07.074

Searching behavior in social Hymenoptera

Schultheiss, P., Cheng, K. and Reynolds, A. M. 2015. Searching behavior in social Hymenoptera. Learning and motivation. 50, pp. 59-67. https://doi.org/10.1016/j.lmot.2014.11.002

Tsallis distributions, Levy walks and correlated-type anomalous diffusion result from state-dependent diffusion

Reynolds, A. M. and Geritz, S. A. H. 2015. Tsallis distributions, Levy walks and correlated-type anomalous diffusion result from state-dependent diffusion. Physica A: Statistical Mechanics and its Applications. 424, pp. 317-321. https://doi.org/10.1016/j.physa.2015.01.034

Venturing beyond the Levy flight foraging hypothesis: reply to comments on 'Liberating Levy walk research from the shackles of optimal foraging'

Reynolds, A. M. 2015. Venturing beyond the Levy flight foraging hypothesis: reply to comments on 'Liberating Levy walk research from the shackles of optimal foraging'. Physics of Life Reviews. 14, pp. 115-119. https://doi.org/10.1016/j.plrev.2015.07.013

Beyond optimal searching: recent developments in the modelling of animal movement patterns as Levy walks

Reynolds, A. M. 2015. Beyond optimal searching: recent developments in the modelling of animal movement patterns as Levy walks. in: Lewis, M. A., Petrovskii, S. V. and Maini, P. K. (ed.) Dispersal, individual movement and spatial ecology. Lecture Notes in Mathematics 2071 Springer, Berlin. pp. 53-76

Pelagic seabird flight patterns are consistent with a reliance on olfactory maps for oceanic navigation

Reynolds, A. M., Cecere. J. G., Paiva, V. H., Ramos, J. A. and Focardi, S. 2015. Pelagic seabird flight patterns are consistent with a reliance on olfactory maps for oceanic navigation. Proceedings of the Royal Society of London Series B Biological Sciences. 282, p. 20150468. https://doi.org/10.1098/rspb.2015.0468

Evidence for a pervasive 'idling-mode' activity template in flying and pedestrian insects

Reynolds, A. M., Jones, H. B. C., Hill, J. K., Pearson, A. J., Wilson, K., Wolf, S., Lim, K. S., Reynolds, D. R. and Chapman, J. W. 2015. Evidence for a pervasive 'idling-mode' activity template in flying and pedestrian insects. Royal Society Open Science. 2, p. 150085. https://doi.org/10.1098/rsos.150085

A priority-based queuing process explanation for scale-free foraging behaviours

Reynolds, A. M., Ropert-Coudert, Y., Kato, A., Chiaradia, A. and Macintosh, A. J. J. 2015. A priority-based queuing process explanation for scale-free foraging behaviours. Animal Behaviour. 108, pp. 67-71. https://doi.org/10.1016/j.anbehav.2015.07.022

Extending Levy search theory from one to higher dimensions: Levy walking favours the blind

Reynolds, A. M. 2015. Extending Levy search theory from one to higher dimensions: Levy walking favours the blind. Proceedings of the Royal Society A: Mathematical, Physical and Engineering Sciences. 471, p. 20150123. https://doi.org/10.1098/rspa.2015.0123

Liberating Levy walk research from the shackles of optimal foraging

Reynolds, A. M. 2015. Liberating Levy walk research from the shackles of optimal foraging. Physics of Life Reviews. 14, pp. 59-83. https://doi.org/10.1016/j.plrev.2015.03.002

Levy flight movement patterns in marine predators may derive from turbulence cues

Reynolds, A. M. 2014. Levy flight movement patterns in marine predators may derive from turbulence cues. Proceedings of the Royal Society A: Mathematical, Physical and Engineering Sciences. 470 (2171), p. 20140408. https://doi.org/10.1098/rspa.2014.0408

Non-volant modes of migration in terrestrial arthropods

Reynolds, D. R., Reynolds, A. M. and Chapman, J. W. 2014. Non-volant modes of migration in terrestrial arthropods. Animal Migration. 2, pp. 8-28. https://doi.org/10.2478/ami-2014-0002

Mussels realize Weierstrassian Levy walks as composite correlated random walks

Reynolds, A. M. 2014. Mussels realize Weierstrassian Levy walks as composite correlated random walks. Scientific Reports. 4, p. 4409. https://doi.org/10.1038/srep04409

Detecting Levy walks without turn designation

Reynolds, A. M. 2014. Detecting Levy walks without turn designation. Behavioral Ecology and Sociobiology. 68, pp. 1893-1899. https://doi.org/10.1007/s00265-014-1819-3

Hierarchical random walks in trace fossils and the origin of optimal search behavior

Sims, D. W., Reynolds, A. M., Humphries, N. E., Southall, E. J., Wearmouth, V. J., Metcalfe, B. and Twitchett, R. J. 2014. Hierarchical random walks in trace fossils and the origin of optimal search behavior. Proceedings of the National Academy of Sciences. 111, pp. 11073-11078. https://doi.org/10.1073/pnas.1405966111

Scaling laws of ambush predator 'waiting' behaviour are tuned to a common ecology

Wearmouth, V. J., Mchugh, M. J., Humphries, N. E., Naegelen, A., Ahmed, M. Z., Southall, E. J., Reynolds, A. M. and Sims, D. W. 2014. Scaling laws of ambush predator 'waiting' behaviour are tuned to a common ecology. Proceedings of the Royal Society of London Series B Biological Sciences. 281, p. 20132997. https://doi.org/10.1098/rspb.2013.2997

Towards a mechanistic framework that explains correlated random walk behaviour: correlated random walkers can optimize their fitness when foraging under the risk of predation

Reynolds, A. M. 2014. Towards a mechanistic framework that explains correlated random walk behaviour: correlated random walkers can optimize their fitness when foraging under the risk of predation. Ecological Complexity. 19, pp. 18-22. https://doi.org/10.1016/j.ecocom.2014.02.004

Distinguishing between Levy walks and strong alternative models: reply

Reynolds, A. M. 2014. Distinguishing between Levy walks and strong alternative models: reply. Ecology. 95 (4), pp. 1109-1112. https://doi.org/10.1890/13-2212.1

Does the Australian desert ant Melophorus bagoti approximate a Levy search by an intrinsic bi-modal walk?

Reynolds, A. M., Schultheiss, P. and Cheng, K. 2014. Does the Australian desert ant Melophorus bagoti approximate a Levy search by an intrinsic bi-modal walk? Journal of Theoretical Biology. 340, pp. 17-22. https://doi.org/10.1016/j.jtbi.2013.09.006

The ontogeny of bumblebee flight trajectories: from naive explorers to experienced foragers

Osborne, J. L., Smith, A. D., Clark, S. J., Reynolds, D. R., Barron, M. C., Lim, K. S. and Reynolds, A. M. 2013. The ontogeny of bumblebee flight trajectories: from naive explorers to experienced foragers. PLOS ONE. 8 (11), p. e78681. https://doi.org/10.1371/journal.pone.0078681

Movement patterns of Tenebrio beetles demonstrate empirically that correlated-random-walks have similitude with a Levy walk

Reynolds, A. M., Lepretre, L. and Bohan, D. A. 2013. Movement patterns of Tenebrio beetles demonstrate empirically that correlated-random-walks have similitude with a Levy walk. Scientific Reports. 3, p. 3158. https://doi.org/10.1038/srep03158

Unravelling the mechanisms of trapline foraging in bees

Liboreau, M., Raine, N. E., Reynolds, A. M., Stelzer, R. J., Lim, K. S., Smith, A. D., Osborne, J. L. and Chittka, L. 2013. Unravelling the mechanisms of trapline foraging in bees. Communicative and Integrative Biology. 6, p. e22701. https://doi.org/10.4161/cib.22701

Effective leadership in animal groups where no individual has pertinent information about resource locations: how interactions between leaders and followers can result in Levy walk movement patterns

Reynolds, A. M. 2013. Effective leadership in animal groups where no individual has pertinent information about resource locations: how interactions between leaders and followers can result in Levy walk movement patterns. EPL. 102, p. 18001. https://doi.org/10.1209/0295-5075/102/18001

Are Levy flight patterns derived from the Weber-Fechner law in distance estimation?

Reynolds, A. M., Schultheiss, P. and Cheng, K. 2013. Are Levy flight patterns derived from the Weber-Fechner law in distance estimation? Behavioral Ecology and Sociobiology. 67, pp. 1219-1226. https://doi.org/10.1007/s00265-013-1549-y

Selection pressures give composite correlated random walks Levy walk characteristics

Reynolds, A. M. 2013. Selection pressures give composite correlated random walks Levy walk characteristics. Journal of Theoretical Biology. 332, pp. 117-122. https://doi.org/10.1016/j.jtbi.2013.04.035

A simple iterative model accurately captures complex trapline formation by bumblebees across spatial scales and flower arrangements

Reynolds, A. M., Lihoreau, M. and Chittka, L. 2013. A simple iterative model accurately captures complex trapline formation by bumblebees across spatial scales and flower arrangements. PLOS Computational Biology. 9, p. e1002938 (10pp). https://doi.org/10.1371/journal.pcbi.1002938

Beating the odds in the aerial lottery: passive dispersers select conditions at takeoff that maximize their expected fitness on landing

Reynolds, A. M. 2013. Beating the odds in the aerial lottery: passive dispersers select conditions at takeoff that maximize their expected fitness on landing. The American Naturalist. 181, pp. 555-561. https://doi.org/10.1086/669677

Radar tracking and motion-sensitive cameras on flowers reveal the development of pollinator multi-destination routes over large spatial scales

Lihoreau, M., Raine, N. E., Reynolds, A. M., Stelzer, R. J., Lim, K. S., Smith, A. D., Osborne, J. L. and Chittka, L. 2012. Radar tracking and motion-sensitive cameras on flowers reveal the development of pollinator multi-destination routes over large spatial scales. PLOS Biology. 10, p. e1001392. https://doi.org/10.1371/journal.pbio.1001392

Gusts within plant canopies are extreme value processes

Reynolds, A. M. 2012. Gusts within plant canopies are extreme value processes. Physica A: Statistical Mechanics and its Applications. 391 (21), pp. 5059-5063. https://doi.org/10.1016/j.physa.2012.05.062

Incorporating sweeps and ejections into Lagrangian stochastic models of spore trajectories within plant canopy turbulence: modeled contact distributions are heavy-tailed

Reynolds, A. M. 2012. Incorporating sweeps and ejections into Lagrangian stochastic models of spore trajectories within plant canopy turbulence: modeled contact distributions are heavy-tailed. Phytopathology. 102, pp. 1026-1033. https://doi.org/10.1094/PHYTO-01-12-0002

Distinguishing between Levy walks and strong alternative models

Reynolds, A. M. 2012. Distinguishing between Levy walks and strong alternative models. Ecology. 93 (5), pp. 1228-1233. https://doi.org/10.1890/11-1815.1

Fitness-maximizing foragers can use information about patch quality to decide how to search for and within patches: optimal Levy walk searching patterns from optimal foraging theory

Reynolds, A. M. 2012. Fitness-maximizing foragers can use information about patch quality to decide how to search for and within patches: optimal Levy walk searching patterns from optimal foraging theory. Journal of the Royal Society Interface. 9, pp. 1568-1575. https://doi.org/10.1098/rsif.2011.0815

Olfactory search behaviour in the wandering albatross is predicted to give rise to Levy flight movement patterns

Reynolds, A. M. 2012. Olfactory search behaviour in the wandering albatross is predicted to give rise to Levy flight movement patterns. Animal Behaviour. 83, pp. 1225-1229. https://doi.org/10.1016/j.anbehav.2012.02.014

Truncated Levy walks are expected beyond the scale of data collection when correlated random walks embody observed movement patterns

Reynolds, A. M. 2012. Truncated Levy walks are expected beyond the scale of data collection when correlated random walks embody observed movement patterns. Journal of the Royal Society Interface. 9, pp. 528-534. https://doi.org/10.1098/rsif.2011.0363

Development and validation of a Lagrangian probability density function model of horizontally-homogeneous turbulence within and above plant canopies

Reynolds, A. M. 2012. Development and validation of a Lagrangian probability density function model of horizontally-homogeneous turbulence within and above plant canopies. Boundary-Layer Meteorology. 142, pp. 193-205. https://doi.org/10.1007/s10546-011-9666-5

Exponential and power-law contact distributions represent different atmospheric conditions

Reynolds, A. M. 2011. Exponential and power-law contact distributions represent different atmospheric conditions. Phytopathology. 101, pp. 1465-1470. https://doi.org/10.1094/PHYTO-01-11-0001

Animal orientation strategies for movement in flows

Chapman, J. W., Klaassen, R. H. G., Drake, V. A., Fossette, S., Hays, G. C., Metcalfe, J. D., Reynolds, A. M., Reynolds, D. R. and Alerstam, T. 2011. Animal orientation strategies for movement in flows. Current Biology. 21, pp. R861-R870. https://doi.org/10.1016/j.cub.2011.08.014

Flight altitude selection increases orientation performance in high-flying nocturnal insect migrants

Aralimarad, P., Reynolds, A. M., Lim, K. S., Reynolds, D. R. and Chapman, J. W. 2011. Flight altitude selection increases orientation performance in high-flying nocturnal insect migrants. Animal Behaviour. 82, pp. 1221-1225. https://doi.org/10.1016/j.anbehav.2011.09.013

Chemotaxis can provide biological organisms with good solutions to the travelling salesman problem

Reynolds, A. M. 2011. Chemotaxis can provide biological organisms with good solutions to the travelling salesman problem. Physical Review E. 83, p. 052901 (4 pp). https://doi.org/10.1103/PhysRevE.83.052901

Chemotaxis can take plant-parasitic nematodes to the source of a chemo-attractant via the shortest possible routes

Reynolds, A. M., Dutta, T. K., Curtis, R. H. C., Powers, S. J., Gaur, H. S. and Kerry, B. R. 2011. Chemotaxis can take plant-parasitic nematodes to the source of a chemo-attractant via the shortest possible routes. Journal of the Royal Society Interface. 8, pp. 568-577. https://doi.org/10.1098/rsif.2010.0417

On the origin of bursts and heavy tails in animal dynamics

Reynolds, A. M. 2011. On the origin of bursts and heavy tails in animal dynamics. Physica A: Statistical Mechanics and its Applications. 390, pp. 245-249. https://doi.org/10.1016/j.physa.2010.09.020

Orientation cues for high-flying nocturnal insect migrants: do turbulence-induced temperature and velocity fluctuations indicate the mean wind flow?

Reynolds, A. M., Reynolds, D. R., Smith, A. D. and Chapman, J. W. 2010. Orientation cues for high-flying nocturnal insect migrants: do turbulence-induced temperature and velocity fluctuations indicate the mean wind flow? PLOS ONE. 5, p. e15758 (6pp). https://doi.org/10.1371/journal.pone.0015758

Bridging the gulf between correlated random walks and Levy walks: autocorrelation as a source of Levy walk movement patterns

Reynolds, A. M. 2010. Bridging the gulf between correlated random walks and Levy walks: autocorrelation as a source of Levy walk movement patterns. Journal of the Royal Society Interface. 7, pp. 1753-1758. https://doi.org/10.1098/rsif.2010.0292

Balancing the competing demands of harvesting and safety from predation: Levy walk searches outperform composite Brownian walk searches but only when foraging under the risk of predation

Reynolds, A. M. 2010. Balancing the competing demands of harvesting and safety from predation: Levy walk searches outperform composite Brownian walk searches but only when foraging under the risk of predation. Physica A: Statistical Mechanics and its Applications. 389, pp. 4740-4746. https://doi.org/10.1016/j.physa.2010.06.027

Maze-solving by chemotaxis

Reynolds, A. M. 2010. Maze-solving by chemotaxis. Physical Review E. 81, p. 062901. https://doi.org/10.1103/PhysRevE.81.062901

Animals that randomly reorient at cues left by correlated random walkers do the Levy walk

Reynolds, A. M. 2010. Animals that randomly reorient at cues left by correlated random walkers do the Levy walk. The American Naturalist. 175, pp. 607-613. https://doi.org/10.1086/651617

A single wind-mediated mechanism explains high-altitude 'non-goal oriented' headings and layering of nocturnally migrating insects

Reynolds, A. M., Reynolds, D. R., Smith, A. D. and Chapman, J. W. 2010. A single wind-mediated mechanism explains high-altitude 'non-goal oriented' headings and layering of nocturnally migrating insects. Proceedings of the Royal Society of London Series B Biological Sciences. 277, pp. 765-772. https://doi.org/10.1098/rspb.2009.1221

Can spontaneous cell movements be modelled as Levy walks?

Reynolds, A. M. 2010. Can spontaneous cell movements be modelled as Levy walks? Physica A: Statistical Mechanics and its Applications. 389, pp. 273-277. https://doi.org/10.1016/j.physa.2009.09.027

Honeybees use a Levy flight search strategy and odour-mediated anemotaxis to relocate food sources

Reynolds, A. M., Swain, J. L., Smith, A. D., Martin, A. P. and Osborne, J. L. 2009. Honeybees use a Levy flight search strategy and odour-mediated anemotaxis to relocate food sources. Behavioral Ecology and Sociobiology. 64, pp. 115-123. https://doi.org/10.1007/s00265-009-0826-2

Levy flight patterns are predicted to be an emergent property of a bumblebee's foraging strategy

Reynolds, A. M. 2009. Levy flight patterns are predicted to be an emergent property of a bumblebee's foraging strategy. Behavioral Ecology and Sociobiology. 64, pp. 19-23. https://doi.org/10.1007/s00265-009-0813-7

Scale-free animal movement patterns: Levy walks outperform fractional Brownian motions and fractional Levy motions in random search scenarios

Reynolds, A. M. 2009. Scale-free animal movement patterns: Levy walks outperform fractional Brownian motions and fractional Levy motions in random search scenarios. Journal of Physics A-Mathematical and Theoretical. 42, p. 434006 (11pp). https://doi.org/10.1088/1751-8113/42/43/434006

Optimising the success of random destructive searches: Levy walks can outperform ballistic motions

Reynolds, A. M. and Bartumeus, F. 2009. Optimising the success of random destructive searches: Levy walks can outperform ballistic motions. Journal of Theoretical Biology. 260, pp. 98-103. https://doi.org/10.1016/j.jtbi.2009.05.033

The Levy flight paradigm - random search patterns and mechanisms

Reynolds, A. M. and Rhodes, C. J. 2009. The Levy flight paradigm - random search patterns and mechanisms. Ecology. 90 (4), pp. 877-887. https://doi.org/10.1890/08-0153.1

Adaptive Levy walks can outperform composite Brownian walks in non-destructive random searching scenarios

Reynolds, A. M. 2009. Adaptive Levy walks can outperform composite Brownian walks in non-destructive random searching scenarios. Physica A: Statistical Mechanics and its Applications. 388, pp. 561-564. https://doi.org/10.1016/j.physa.2008.11.007

Predator percolation, insect outbreaks and phase polyphenism

Reynolds, A. M., Sword, G. A., Simpson, S. J. and Reynolds, D. R. 2009. Predator percolation, insect outbreaks and phase polyphenism. Current Biology. 19, pp. 20-24. https://doi.org/10.1016/j.cub.2008.10.070

Does a 'turbophoretic' effect account for layer concentrations of insects migrating in the stable night-time atmosphere?

Reynolds, A. M., Reynolds, D. R. and Riley, J. R. 2009. Does a 'turbophoretic' effect account for layer concentrations of insects migrating in the stable night-time atmosphere? Journal of the Royal Society Interface. 6, pp. 87-95. https://doi.org/10.1098/rsif.2008.0173

Aphid aerial density profiles are consistent with turbulent advection amplifying flight behaviours: abandoning the epithet 'passive'

Reynolds, A. M. and Reynolds, D. R. 2009. Aphid aerial density profiles are consistent with turbulent advection amplifying flight behaviours: abandoning the epithet 'passive'. Proceedings of the Royal Society of London Series B Biological Sciences. 276, pp. 137-143. https://doi.org/10.1098/rspb.2008.0880

Bee searching strategies in the landscape

Osborne, J. L., Smith, A. D. and Reynolds, A. M. 2008. Bee searching strategies in the landscape. Abstracts 3rd European Conference of Apidology (EurBee3), Belfast,8-11 September 2008 . pp. 79

How many animals really do the Levy walk? Comment

Reynolds, A. M. 2008. How many animals really do the Levy walk? Comment. Ecology. 89 (8), pp. 2347-2351. https://doi.org/10.1890/07-1688.1

Deterministic walks with inverse-square power-law scaling are an emergent property of predators that use chemotaxis to locate randomly distributed prey

Reynolds, A. M. 2008. Deterministic walks with inverse-square power-law scaling are an emergent property of predators that use chemotaxis to locate randomly distributed prey. Physical Review E. 78, p. 011906 (5 pages). https://doi.org/10.1103/PhysRevE.78.011906

Particle deposition onto rough surfaces

Lo Iacono, G., Reynolds, A. M. and Tucker, P. G. 2008. Particle deposition onto rough surfaces. Journal of Fluids Engineering. 130, p. 074501 (5 pages). https://doi.org/10.1115/1.2948359

Avoidance of conspecific odour by carabid beetles: a mechanism for the emergence of scale-free searching patterns

Guy, A. G., Bohan, D. A., Powers, S. J. and Reynolds, A. M. 2008. Avoidance of conspecific odour by carabid beetles: a mechanism for the emergence of scale-free searching patterns. Animal Behaviour. 76, pp. 585-591. https://doi.org/10.1016/j.anbehav.2008.04.004

Optimal random Levy-loop searching: new insights into the searching behaviours of central-place foragers

Reynolds, A. M. 2008. Optimal random Levy-loop searching: new insights into the searching behaviours of central-place foragers. EPL-Europhysics Letters. 82, p. 20001 (6pp). https://doi.org/10.1209/0295-5075/82/20001

Modelling of concentrations along a moving observer in an inhomogeneous plume. Biological application: model of odour-mediated insect flights

Lo Iacono, G. and Reynolds, A. M. 2008. Modelling of concentrations along a moving observer in an inhomogeneous plume. Biological application: model of odour-mediated insect flights. Environmental Fluid Mechanics. 8, pp. 147-168. https://doi.org/10.1007/s10652-008-9054-y

The influence of search strategies and homogeneous isotropic turbulence on planktonic contact rates

Rhodes, C. J. and Reynolds, A. M. 2007. The influence of search strategies and homogeneous isotropic turbulence on planktonic contact rates. EPL-Europhysics Letters. 80, p. 60003 (5pp). https://doi.org/10.1209/0295-5075/80/60003

Honeybees perform optimal scale-free searching flights when attempting to locate a food source

Reynolds, A. M., Smith, A. D., Reynolds, D. R., Carreck, N. L. and Osborne, J. L. 2007. Honeybees perform optimal scale-free searching flights when attempting to locate a food source. Journal Of Experimental Biology. 210, pp. 3763-3770. https://doi.org/10.1242/jeb.009563

Avoidance of conspecific odour trails results in scale-free movement patterns and the execution of an optimal searching strategy

Reynolds, A. M. 2007. Avoidance of conspecific odour trails results in scale-free movement patterns and the execution of an optimal searching strategy. EPL-Europhysics Letters. 79, p. 30006 (5 pages). https://doi.org/10.1209/0295-5075/79/30006

Dissipation conditioned stochastic modeling of scalar concentration fluctuations in turbulent flows

Reynolds, A. M. 2007. Dissipation conditioned stochastic modeling of scalar concentration fluctuations in turbulent flows. Physics Of Fluids. 19, p. 075101 (6pp). https://doi.org/10.1063/1.2747681

Displaced honey bees perform optimal scale-free search flights

Reynolds, A. M., Smith, A. D., Menzel, R., Greggers, U., Reynolds, D. R. and Riley, J. R. 2007. Displaced honey bees perform optimal scale-free search flights. Ecology. 88 (8), pp. 1955-1961. https://doi.org/10.1890/06-1916.1

Appetitive flight patterns of male Agrotis segetum moths over landscape scales

Reynolds, A. M., Reynolds, D. R., Smith, A. D., Svensson, G. P. and Lofstedt, C. 2007. Appetitive flight patterns of male Agrotis segetum moths over landscape scales. Journal of Theoretical Biology. 245, pp. 141-149. https://doi.org/10.1016/j.jtbi.2006.10.007

Free-flight odor tracking in Drosophila is consistent with an optimal intermittent scale-free search

Reynolds, A. M. and Frye, M. A. 2007. Free-flight odor tracking in Drosophila is consistent with an optimal intermittent scale-free search. PLOS ONE. 2, p. e354. https://doi.org/10.1371/journal.pone.0000354

Optimal scale-free searching strategies for the location of moving targets: new insights on visually cued mate location behaviour in insects

Reynolds, A. M. 2006. Optimal scale-free searching strategies for the location of moving targets: new insights on visually cued mate location behaviour in insects. Physics Letters A. 360, pp. 224-227. https://doi.org/10.1016/j.physleta.2006.08.047

Ballooning dispersal in arthropod taxa with convergent behaviours: dynamic properties of ballooning silk in turbulent flows

Reynolds, A. M., Bohan, D. A. and Bell, J. R. 2006. Ballooning dispersal in arthropod taxa with convergent behaviours: dynamic properties of ballooning silk in turbulent flows. Biology Letters. 2, pp. 371-373. https://doi.org/10.1098/rsbl.2006.0486

Stochastic simulation of Lagrangian trajectories in near-wall turbulence

Reynolds, A. M. 2006. Stochastic simulation of Lagrangian trajectories in near-wall turbulence. Physics Of Fluids. 18, p. Art no. 075107. https://doi.org/10.1063/1.2236303

On the intermittent behaviour of foraging animals

Reynolds, A. M. 2006. On the intermittent behaviour of foraging animals. EPL-Europhysics Letters. 75, pp. 517-520. https://doi.org/10.1209/epl/i2006-10157-x

On chirality and turbulent dispersion

Reynolds, A. M. 2006. On chirality and turbulent dispersion. Physica D-Nonlinear Phenomena. 218, pp. 185-190. https://doi.org/10.1016/j.physd.2006.05.008

Cooperative random Levy flight searches and the flight patterns of honeybees

Reynolds, A. M. 2006. Cooperative random Levy flight searches and the flight patterns of honeybees. Physics Letters A. 354, pp. 384-388. https://doi.org/10.1016/j.physleta.2006.01.086

On the anomalous diffusion characteristics of membrane-bound proteins

Reynolds, A. M. 2005. On the anomalous diffusion characteristics of membrane-bound proteins. Physics Letters A. 342 (5-6), pp. 439-442. https://doi.org/10.1016/j.physleta.2005.05.086

Parameterizations of Lagrangian spin statistics and particle dispersion in the presence of coherent vortices

Veneziani, M., Griffa, A., Reynolds, A. M., Garraffo, Z. D. and Chassignet, E. P. 2005. Parameterizations of Lagrangian spin statistics and particle dispersion in the presence of coherent vortices. Journal Of Marine Research. 63, pp. 1057-1083. https://doi.org/10.1357/002224005775247571

Scale-free movement patterns arising from olfactory-driven foraging

Reynolds, A. M. 2005. Scale-free movement patterns arising from olfactory-driven foraging. Physical Review E. 72, p. Art. no. 041928 (5 pages). https://doi.org/10.1103/PhysRevE.72.041928

A Lagrangian stochastic model for the dispersion and deposition of Brownian particles in the presence of a temperature gradient

Lo Iacono, G. and Reynolds, A. M. 2005. A Lagrangian stochastic model for the dispersion and deposition of Brownian particles in the presence of a temperature gradient. Journal Of Aerosol Science. 36, pp. 1238-1250.

Predictions for particle deposition from LES of ribbed channel flow

Lo Iacono, G., Tucker, P. G. and Reynolds, A. M. 2005. Predictions for particle deposition from LES of ribbed channel flow. International Journal Of Heat And Fluid Flow. 26, pp. 558-568.

On the distribution of Lagrangian accelerations in turbulent flows

Reynolds, A. M., Mordant, N., Crawford, A. M. and Bodenschatz, E. 2005. On the distribution of Lagrangian accelerations in turbulent flows. New Journal Of Physics. 7, p. Art. no. 58 (9 pages).

Derivation of a Fokker-Planck equation for generalized Langevin dynamics

Khan, S. and Reynolds, A. M. 2005. Derivation of a Fokker-Planck equation for generalized Langevin dynamics. Physica A: Statistical Mechanics and its Applications. 350, pp. 183-188.

Stochastic modeling of protein motions within cell membranes

Khan, S., Reynolds, A. M., Morrison, I. E. G. and Cherry, R. J. 2005. Stochastic modeling of protein motions within cell membranes. Physical Review E. 71, p. Art. no. 041915 (7 pages). https://doi.org/10.1103/PhysRevE.71.041915

On the simulation of particle trajectories in turbulent flows

Reynolds, A. M. and Lo Iacono, G. 2004. On the simulation of particle trajectories in turbulent flows. Physics Of Fluids. 16 (12), pp. 4353-4358. https://doi.org/10.1063/1.1804551

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